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The slanted e is a callback to the previous Google logo, which also had a slanted e.

The O's were slightly tilted too, no?

They still are in the new logo. They're perfect circles, but they've been rotated to the exact same angle as the old "o"s.

Made me look!

Screenshotting the doodle, the counters are 24×26 pixels. Not perfect circles, and not tilted.

how do you rotate a perfect circle

that's... the joke.

By turning it so many degrees.