>What can freemium webapps learn from freemium games?
Games have some advantages that can be hard for apps to replicate; games are, in general, more social than typical applications. Games are also somewhat more enviable. When I was 11, a few friends and I played RuneScape (remember runescape?). After one of us got a paid membership, everyone else suddenly got way more interested in having a paid RuneScape membership. That is to say, if one person gets a paid membership to a multiplayer online game, other people in their social circle will probably be more likely to subscribe as well.
Taking advantage of this with a webapp would require that you provide something that someone either finds worthwhile showing off or something conspicuous enough that they don't have to. The second condition applied for Apple's iPod in the early part of the decade. It'd also probably be a good idea to offer benefits to groups of paid members as well as to individual paid members.