Yes. It gossips with the following list of servers. Keys are uploaded and also fetched form these servers:
'https://pgp.mit.edu', 'http://pool.sks-keyservers.net', 'http://keys.gnupg.net', 'http://keyserver.ubuntu.com', 'http://pks.gpg.cz'
'https://pgp.mit.edu', 'http://pool.sks-keyservers.net', 'http://keys.gnupg.net', 'http://keyserver.ubuntu.com', 'http://pks.gpg.cz'