They hit an issue no-one really thinks about: resizing hash tables. No-one thinks about it, as it happens transparently under the surface, but for them, the few milliseconds it took to resize the hash tables would have enormously detrimental effects on just about every other Google service. The solutions they were talking about were really quite amazing. That's the insanity they face.
So, if anyone hasn't seen Google Sparsehash[1], check it out: it's another near drop-in improvement to the STL. Two bits per entry of overhead and insanely fast. I can't remember if the Quota team used it specifically, but I wouldn't be surprised if it's one of the results of their work.