They didn't leave the architecture alone, right? They shoved React in and created a weird SSR + React frankenstein that is objectively worse in many ways.
It's just pointless hacks on hacks. GitHub didn't need React on the frontend and any potential resource savings of client side rendering were lost when they realized they need to do SSR on that stuff too.
I've encountered so much frontend jank as they expanded that portion of the stack whereas it was always excellent when it was just Ruby SSR and minimal JS on the frontend.