- Vertical tabs
- Drag & Drop between "sticky" and "ephemeral" sections
- Hiearchically nested folders
- Command palette with fuzzy match to open/recent tabs
- Meeting integration
There were some other nice features ("Smart" folders with GH PRs come to mind), but really 95% of my usage was due to the above. The tab model was so good it actually allowed me to replace "to do list" type tools with just folders of tabs.
For me, I think they had their target market all wrong. They built a bunch of design features nobody used and cared about (custom theming?), when the core pitch was very simple: People LIVE in their web browsers, but everyone just seems to accept that having 100 impossible-to-read chrome tabs open at the same time is somehow acceptable. There are so many browser "power users" out there in white collar roles that don't even realize a better world is possible. These people are not going to be downloading (and likely can't download) a sketchy sounding "firefox fork" named "Zen", but they will absolutely download a professional-looking Chrome fork with security guarantees that allows them to organize their work better.