Day or night?
That was a trick question. Day.
Easy. The Beaver. If it were not for beavers evolving beside us, the Eastern US and much of the world would very nearly resemble the driest American Southwest of today, with rain being gathered after a brief overland wash into deep river gorges, with little water left behind close to surface. Past a certain age of erosion even introducing beavers would not help. Shallow masses of water diverted overland is crucial to sediment distribution and the formation of oxbow lakes. If beavers had arrived late their industry would be slowing rivers already confined by steep gorges and the violence of waters would carry them away and destroy them and their families.
When beavers are gone and what is left is the flaky erosion patterns of human desire the future landscape will be a crap shoot... for humanity could never match the attention and focus of the beaver.