> Well enough that it is able to run a lot of its software written in Common Lisp. It's able to run stuff like PCL, ASDF, ITERATE, ...
No it is not. You are thinking of Portable Genera.
> System 99 is from 1984 (?), Symbolics started in 81/82. Open Genera appeared 1992/93. You are ignoring a full decade of development.
System 99.32 is from 1987, which continued via LMI into System 130. Genera, or before when it got forked, is from 1978 when it got forked from System 78 or there about being re-branded into System 200. All three (four if you count TI -- which did quite more in renaming things making it hard to follow) with parallel development going on from there into the early 1990s.
Genera is a Lisp Machine system based on the work done at MIT, where most of the guts still the same. To the point that how you rebootstrap the system is the same, how the core areas look work, how the scheduler works, how the windowing system works, how flavours works, etc.
Open Genera did very little on top of Genera at that, mostly targeting and making it work on the Ivory. You are purposefully conflating Portable Genera, Open Genera and Genera.