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I still don't understand the title, no matter how hard I try. Is there a word missing? A misspelling?

"How many bathrooms have (fossilized remains of) Neanderthals in the (wall/floor) tile?"

I had to start reading the article before I was able to parse it correctly.

Just got it. Have is the possessive definition, not the helper verb for a past participle.

Read like: How many tiles contain Neanderthals within their tiles?