> using email addresses from previous breach corpuses
> enumerating a publicly accessible resource
> Trello advised that no unauthorised access had occurred
That sounds like a credential stuffing attack to me!
Most likely the attackers used the password reset form, and that Trello acknowledge the existence of emails. This used to be considered bad practice, but it's more nuanced than that, provides basically no useful information, and is now considered to be a relatively safe practice.
Alternatively it's possible they used some sort of sharing functionality to try to share Trello content with other addresses, and again the availability of that for usage would be by-design.