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Do these folks now see single characters or will this adapt?

This uses `rem`s for font sizes, so it will increase if you increase your system font. This is what it looks like if you set 32px as default instead of 16px: https://i.imgur.com/amr8t8Q.png

The three-line “The best place to find government services and information” doesn’t make sense when screen space is scarce due to large fonts.

In addition, old folks tend to have difficulty finding information when they have to scroll down, especially when there’s no indication that there’s more below the fold (e.g. formerly scroll bars).

There is no fold.

https://www.lukew.com/ff/entry.asp?1946

How important is gov.uk's tagline? It seems like making the content larger and on screen is more important than having a HUUUGE tagline.

Perhaps, on mobile. But why should it take so much space on a huge monitor?

great point - i spent seven years in customer facing banking which included helping people with the mobile app. nearly everyone over 50 had large font sizes.