But there are numerous ways to use feature flags incorrectly - typically once you have multiple long-lived flags that interact with each other, you've lost the thread. You no longer have one single application, you have n_flags ^ 2 applications that all behave in subtlety different ways depending on the interaction of the flags.
There's no way around it - you have to test all branches of your code somehow. "Just let the users find the bugs" doesn't work in this case since each user can only test their unique combination of flags. I've regularly seen default and QA tester flag configurations work great, only to have a particular combination fail for customers.
The only solution is setting up a full integration test for every combination of flags. If that sounds tedious (and it is), the solution is to avoid feature flags, not to avoid testing them!