The article also points out that GMs LT6 flat-plane V8 for the Corvette Z06 has a sound different from the F355, this apparently being deliberate design choice made by choosing which harmonics to emphasize in the exhaust system.
You say that flat-plane engines do not require the complex header primaries that have to cross over from one bank to the other, but AFAIK typical road car and truck V8s do not have such headers[1]. Would the purpose of such headers be to give a cross-plane V8 scavenging about as good as a flat-plane one, all else being equal? In that case, would the cross-plane engine lose most of its distinctive sound?
I would be interested in hearing the difference between otherwise comparable flat-plane and cross-plane V8s at the same RPM.