For instance, I enjoy how the same region will evolve over time depending on where you are in your main quest. At first, it's just a road to go to the temple. But as the game progress and you get more items, you can explore it in depth and discover awesome quests/items.
I agree with him/her that there's really few games out there that mimics the game mechanics of zelda. Mario 64 comes close but, for instance, miss the in depth strategy and the various items. Zelda really have unique game play and it's not just about the world of Zelda.. you could create a totally different world and still copy the game mechanics.. but lots of really successful game use only a part of zelda's game play which is on what the author argue.
In fact, it can be summarized as Zelda being the intersection between multiple 'genres'. And it's surprising that nobody copies it exactly as we all know how Zelda is a success and is popular.