This article uses the word 'rune' extensively. From the context I assume it means 'lexem' or 'token' (i.e. the unit the lexer/scanner produces and feeds to parser). But then the article uses the word 'token' to mean the output of a parser ('keyword token'), while the usual terminology is that parser output is called a 'parse tree'.

So, in this terminology, the parser consumes 'runes' and outputs 'tokens', while the usual terminology is that parser consumes 'tokens'/'lexems' and outputs 'parse tree'.