It's answered in the article:
> They argue this reduces the threshold for a so-called 51 percent attack. "If a node operator's self-interest is to be dishonest, then there is no explicit penalty for doing so. Moreover, the number of entities necessary to execute a 51 percent attack on Bitcoin was reduced from 51 percent of the entire network (which we estimate at approximately 59,000 nodes) to only the four most popular mining pool nodes (less than 0.004 percent of the network)," the study found.