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It is federated like email so if you have an email (@gmail, @yahoo, @custom.domain, etc) they are all interoperable. (Modulo moderation / spam filters)

You don't have nor would be able to create an email address in all public email servers. (Without mentioning private/non-public email servers or personal/family ones of others)

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I login to my old Mastadon account on some specific server then try to reply to someone on another one. It asks me for the domain I want to act from so I type it in. Click Proceed to reply and am greeted with an error screen saying something went wrong. I really want an alternative to Twitter, but using this site feels very abrasive.

The reply page is hosted on the remote instance often so they can show anything including an error page. I think qoto.org has some extra changes that make this a little more well behaved by hosting it locally (depending on which button you push.)

looks that there are some issues with the huge wave of new users. I have the same problem trying to reply or follow accounts from another few instances.

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