Almost certainly the ad revenue goes towards the authors of the original song, due to how music licensing works.
David Paich gets paid every time a radio station plays wheezer's cover of Africa, while the wheezer gets no money for radio play of that song.
However, "Surfin' USA" goes to whomever owns Chuck Berry's rights to "Sweet little sixteen" -- those were much more similar than "Born This Way" and "Express Yourself" IMO.
Copyright owners could still put in manual Content ID claims, though.