I would clarify that to say numbers were inseparable from
words for them. Writing has only existed a short time of our history, so numerals are pretty recent.
And the "meaning" of zero as a number like others along a number line, rather than as mere notation for "nothing", I assume only ever became necessary with the invention of negative numbers.
With addition, multiplication and division, zero simply does nothing or annihilates a number, and so doesn't need to be treated like other numbers. AFAIK, zero as a number arises first in figuring out how to "get to" negative numbers, e.g. what is two minus four (one, zero, negative one, negative two), where zero is required as a numeric concept.
Negative numbers were a big step forwards. Zero, I still don't see it -- either it was just convenient notation for "nothing", or part and parcel of the shift to negative numbers. Unless I'm missing something in the historical record?