If it’s replacing an ipsec mesh that’s pretty hard to believe. And if that was the issue and commercial support couldn’t even identify that as the cause, ZeroTier has bigger issues.
Finally, how can ZeroTier’s support be so incompetent to not recognize connectivity issues between endpoints? That’s one of the few things that goes wrong with tunnel meshes.
Symmetric NAT basically breaks everything that doesn't use a simple client/server hub-and-spoke networking model.