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I personally find that screen tries to do less, and therefore, gets in my way less. Also, emacs-style keybindings fit better with the general interaction model of the shell, which also uses emacs-style keybindings.

I know that one can switch to vim-style keybindings in the shell, but they have never felt as "comfortable" (and I use vim as my daily driver).

Last but not least, the default keybinding for splitting in tmux (C-b ") is just awkward. Screen has way better defaults.

TBH I don't think so. Screen is great, but overall tmux is better. Screen is often shipped by default with linux OSes so maybe there's some advantage for airgapped machines, etc.

I have used it in the past to connect to devices over serial.

Familiarity.

Same reason as the reason to use a prius when maseratis are available. sometimes a simple tool is better