My reasoning:
(1) As the article said, full buses move slower because more passengers means more boarding/unboarding time at stops.
(2) The lead bus is the fuller, slower one. Slower bus in front means buses get closer together.
(3) If one bus passes the other, then you have the faster bus in front and the slower one behind. They should then get further apart.
At least they should for a time. The now-leading bus is going to scoop up a lot of passengers at the next several stops and the now-trailing bus won't, so the leading bus will eventually slow down. So it's not perfectly self-balancing or anything, but maybe it would get some distance between them. More distance than not doing it.