"Knuth had previously proved an exponential upper bound on the number of Topswops steps, and conjectured that one might also prove a matching lower bound. What Dr. Hal Sudborough and Dr. Linda Morales did, however, was to prove a lower bound that is much better than that proposed in Knuth’s conjecture..."
However, the paper cited says “A Quadratic Lower Bound for Topswops”, so the lower bound proved is much worse than the one conjectured by Knuth.