You can hardly tie ressource usage and startup time to the number of features a DE has.
Take Windows as an extreme. Slow as all balls, around 1.5 GB RAM usage and essentially no features that go beyond the basics.
On the other hand, take Enlightenment. It's got fancier graphics effects than KDE, nearly as much customization and I'd say about half the features (which is still a lot more than Windows, GNOME, even Xfce),
and it starts up in the blink of an eye, has extremely little lag compared to other DEs, and only uses about 150 MB.
It's also really buggy and not really a complete, polished DE, so despite all its advantages, I would only recommend it, if you really feel comfortable with desktop Linux and don't mind a bit of dicking around to get things working the way you want them to.
And GNOME clocks in at around 1 GB RAM usage, making it the most heavyweight Linux DE, despite it lacking tons of features that pretty much any other DE has.
(KDE nowadays clocks in at around 600 MB, Xfce at around 350 MB.)