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The article ends with: "It doesn’t mean it has an evolutionary basis."

If not evolution, then what? Doesn't all behavior have an evolutionary basis?

No. There is a technical term for this - can someone help me out? I can't remember it....

Yes. They're called non-adaptive traits or spandrels: https://en.wikipedia.org/wiki/Spandrel_(biology)

Philanthropy?

Selflessness?

Sacrifice?