Right. I thought it was significant that the two lead investors are strategic (the first big tranche came from Google,
not GV), so there's even less signaling there.
As for $1.4B vs $500K... they're different species of investment animal. The people making the hundred-million dollar decisions aren't prepared to make hundred-thousand dollar decisions. So I don't think those huge investments are actually replacing or crowding out seed funding (especially if a highly-funded company like ML can then buy a bunch of seed-funded companies, returning liquidity to the seed market).
The only thing that's weird about it is money is fungible, so the two are truly equivalent... just not commutable. Could those billions have been a shower of seed investments instead? I dunno; this is one of those things that makes me scratch my head and shrug. I don't know enough about the history of finance, and venture finance in particular, to weigh in on the logic (or forces of nature, or chance) that led to this structure, but it seems to make sense except when it doesn't. (/hedge)