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we've been on Fleep with our distributed team of a dozen people for a year now and very happy with it. the quickest way to think about them is that if Slack is a modern take of IRC, then Fleep makes Skype's model of persistent p2p chats work (fast, mobile, syncing well, file storage, some clever UI improvements, web access).
The thing that makes emails of great is that it is not a program controlled by a single commercial entity but instead an open standard. Although I find this an interesting prospect, I see the potential for this particular product is quite limited because of the aforementioned plus their pricing scheme.
Of course, this wouldn't be the first time a commercial entity saw that as a bug and not a feature. How many "email must die and $SERVICE will kill it" articles have we seen recently?
I tried to create an account. Still waiting for the activation email so I can "click the link and get started".
Actually tons of services are trying to do so, but interesting to see who wins at the end...