Unless the license agreement for contributing specifically says the author forfeits their copyright, but no open source licenses I know of do this.
Forfeiting copyright is not possible in some jurisdictions.
I'm not sure what is unclear about a refutation of a single sentence.
Moral rights are generally impossible to forfeit and assign. Economic rights usually (I don't know of any jurisdictions where assigning economic rights to the project maintainer would be impossible - are you?) are possible to assign.
In common law systems, the two are usually traditionally much more closely tied, and when we talk about copyright in English, it is usually the economic rights we're referring to.
I know to contribute to Clojure/ClojureScript-proper, I had to sign http://clojure.org/contributing
The short version is assign FSF copyright, or put it in the public domain.