OK, actually, I agree with some of this. I didn't think labeling that behavior as "primal" was accurate or helpful. It's certainly social, and bound to be more or less strong depending on the person and situation. Calling it primal is defeatist, plus it assumes that men have to fight against their base nature to be decent people.
On the other hand, I don't think the article says men don't get rape threats, just that they're rare. Also I think society really doesn't understand how sexist it is.