> It's possible that for it to be published, a higher standard would be needed. That is, the actions they took were ethical, but perhaps inappropriate for scientific publication.
> But, if that were the case, the peer reviewers and the journal in which it was published should have flagged that. That it was published shows they didn't have any significant concerns.
This would not be a scientific ethics issue if they explained their IRB review in the manuscript. It is so unusual to not do this that it is a reasonable assumption something funny is going on, in my experience.
For example, the journal could be incentivized to look the other way in order to publish a high publicity article. I'm NOT saying that's what PNAS did, but just because something is published doesn't mean it's ethical. (See: https://www.ncbi.nlm.nih.gov/pubmed/20137807)