Today, we're keeping everything on SSD -- twice, in fact, to maintain a hot spare for each server. SSD prices have fallen to the point where, even renting from Amazon, we can make the cost model work; and SSD is a great crutch as we tune our storage layer. But spinning disk will be a lot cheaper.
As for Java: we've been pretty surprised ourselves. Going in, we expected to need a lot more "native" (C) code. As it turned out, currently the only non-Java code we're using is the LZ4 library, and even there we only recently moved to the non-Java implementation. We do soon expect to move a few more bits of code to C, such as the core text-search loop. But based on the performance we've been getting in pure Java, we don't anticipate any need to move more than a fraction of 1% of our overall code base.
We do have some code that lives in a sort of middle ground -- Java code written like it was in C. Our database buffers live in a set of 2GB byte arrays, soon to be replaced by native memory blocks. We do our own (very simple) memory allocation within that, and use a sprinkling of Unsafe operations to access it. This gets us closer to C performance (not all the way there), without any of the hassles of bridging across languages. This part is still well under 1% of our overall codebase.