After reading this blog post, I think the rule of thumb should be "If I take a screenshot of your app at any moment (except during animations), it must make sense". I don’t think making sense during an animation should really be a goal, as long as it makes sense before and after.
In an ideal world, it is hard to argue with. Yes, sure it should make sense. But also, please don't spend precious cycles on this unless all the other bugs are fixed, and this animation consistency is truly the most important remaining issue to address.
Maybe I've just spent too many years as a pixel-perfect chasing frontend developer, but things can look very janky if they jump out of place during animations, compared to where they are before/after.
My comment starts with "After reading this blog post"; of course I read it, but it left me totally unconvinced, especially because the author doesn’t bother showing good examples: he criticizes these and then leave you with a random raccoon animation and that’s all. For example, I don’t understand what’s wrong with the Youtube animation; it looks perfectly fine to me.