The simulation is so simplified that I see no terms for the control of pedaling. Riding a real bicycle isn't just about steering and leaning a bit. You need to propel the bicycle a certain amount.
The paper buries this in the following:
>Although the two-neuron network controller works well for a range of speeds, one thing the controller does not do is to try to dampen the instabilities that can arise when riding too slowly or in too sharp of a turn. (This would probably require a third neuron that isdedicated to this task.)
They say 'damping instabilities' but it is way more than that, because as anyone who has learned to ride a bike knows, the hard part is getting started at that zero point of forward velocity - how to apply torque to the crank at the same time as compensating with the steering to balance at such low momentum. It's not a trivial solution to 'damping instabilities' when getting going in the first place is the most difficult part (as any 5 year old child will demonstrate).