By itself it's not a tell but combined with all else it's hard to pass by. Author's other article from 2025 has less than half the dashes and it's the same length
I haven't found a satisfactory explanation, but whatever the explanation is, it is undoubtedly true that LLMs use them to an almost absurd extent compared to the vast majority of human writers. Anyone who reads a lot of prose can see that.
And they don't have a sense of stylistic restraint, so they often go overboard with one or more of the above.
I don't know about the structures you mention specifically but if you compare an article on humanities or social matters against the style that's common in science and technical writing, chances are it's going to look more verbose in any case.
I don't necessarily have the best AI-dar but TFA didn't ring any LLM bells to me.