... just speaking as someone who sometimes has to review very long PRs sometimes, though, I feel like 25% is a roughly normal level of "signal to noise." 5,000 lines of core logic is a LOT, and the tests and dependencies do still need to be read.
EDIT: I feel like the problem, as a reviewer, is processing 4 months of intensive research/development and providing useful feedback. At that point, there's probably not much major input you can have into the core architecture or strategy, so you're probably not providing much more than a bugbot at that point.