Which mainboards are cheap and have 4 pcie16x (electrical) slots, that don't need weird risers to fit 4 GPUs
What's cheap to you? You can find Epyc 7002/7003 boards on ebay in the $400 range and those will do it. That's probably the best deal for 4x PCIE 4.0 x16 and DDR4. Probably $500 range with a CPU. That's in the ballpark of a mid to high end consumer setup these days.
If I had more income, I would also buy 4x 96g Optane drives of p0 swap disks and a few ssd's for p1 swap disks. To evaluate how well you can get a 1T model running in these absurd ram prices.
What I eventually had to do was buy a used Threadripper box to run those cards, because PCIe Gen 4 definitely works.