I do wonder if those animals have things like valves in their veins, as I understand it if the circulatory system wasn't as complex as it is, heart would have to pump a lot harder to move the volume it does. This isn't an area I know much of anything about, I just know veins have valves and can expand and contract to different stimuli much as a heart can... so even though mammals have one heart it's not like the rest of the system is a static not helping to pump blood.