Because offhand, I know you could do things like cute optimizations of redundant data to minimize seek time on optical media, but with HDDs, you get no promises about layout to optimize around...
The only thing I can think of is if it was literally something as inane as checking the "store deduplicated by hash" option in the build, on a tree with copies of assets scattered everywhere, and it was just nobody had ever checked if the fear around the option was based on outcomes.
(I know they said in the original blog post that it was based around fears of client performance impact, but the whole reason I'm staring at that is that if it's just a deduplication table at storage time, the client shouldn't...care? It's not writing to the game data archives, it's just looking stuff up either way...)