A lot of that group is making use of the IO capabilities of these systems to run lots of PCI-E devices & hard drives. There's not exactly a cost-effective modern equivalent for that. If there were cost-effective ways to do something like take a PCI-E 5.0 x2 and turn it into a PCI-E 3.0 x8 that'd be incredible, but there isn't really. So raw PCI-E lane count is significant if you want cheap networking gear or HBAs or whatever, and raw PCI-E lane count is $$$$ if you're buying new.
Also these old systems mean cheap RAM in large, large capacities. Like 128GB RAM to make ZFS or VMs purr is much cheaper to do on these used systems than anything modern.