https://en.wikipedia.org/wiki/LORAN
Before LORAN, used radio beacon navigation and star navigation (from Newton time), and good human navigator could achieve about 50km precision.
You could easy see signs of star navigation on good preserved old planes - they all have some sort of fully glass dome, or blister, to provide good near semi-sphere view. And sure, all those before-GPS era planes have separate navigator job position, sometimes shared with mechanic.
https://www.reddit.com/r/WWIIplanes/comments/59xfkz/pby_wais...
You could ask, how planes could fly with 50km precision? Answer is easy - at all plane routes built ground structures easy seen from air and last mile navigation become essentially visual flight, nothing more, nothing less.
On some places ground navigation structures preserved now, for examples:
https://en.wikipedia.org/wiki/Airway_beacon
https://en.wikipedia.org/wiki/Transcontinental_Airway_System
Perhaps in general aviation, but I can't think of any modern commercial airliner without an INS via the air data inertial reference unit.
The INS unit is separate and often has its own set of gyros and has to be connected as separate input to FMS or other navigational computers, same as connecting GPSes or other radio nav components.
For example the current model of popular Universal Avionics UNS1 series of NCU (navigational computer part of FMS) come with built-in augmented GPS receiversz but do not mention INS functionality at all even in extended models. Don't have access to manuals at the moment, but I'd expect to see INS as optional to connect over one of the external connectors on the MCU, as it was on the older models without integrated GPS
I had assumed that the ADIRU’s inertial reference data from its gyroscopes and accelerometers would feed into the FMS to provide INS capabilities in case GNSS was unavailable.
GNSS becoming "reasonably easy" to add meant that there was way less push to integrate INS after early use in transoceanic flights