Surely that's a reference to the biologist mentioned in another comment, who said God had "an inordinate fondness of beetles".
Pratchetts's books are full of that kind of reference.
Pratchetts's books are full of that kind of reference.
But then he was talking about the "pinnical" of life and evolution. Ponder thought he meant humans, capable of mastering their world.
Then he looked down and saw the leg twitch...