I did not read the paper, so I cannot comment on the "solid grasp of class inclusion", but regarding the capacity that you described in your comment, I have a 2-year-old and it's been a long while since she has mastered this (book vs this book, toy vs toy, fruit vs an apple and so on). As far as I know, most two year old have already acquired this concept.
(EDIT I see the other comment says something similar and you have replied)