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if a protomolecule ever comes to be, it's going to be some sort of fungi-based lifeform.

Is that an Expanse reference or does the word mean something else in biology?

It definitely doesn't mean anything in real science.

it is! but I meant the idea of it, ie self-replicating biological programs that could be hurled through space and be used for terraforming.

Tell me about it ....