Two reliable effects predict runaway complexity for any initially simple life form in a non-trivial environment.
1) BOOT STRAPPING COMPLEXITY: Non-trivial static environment: Something simple is rarely the global efficiency optimum in a non-trivial environment. There is nothing trivial about chemistry and the myriad of terrains created by physics in the non-living world. So simple living things, in competition, quickly get more complex.
2) ACCELERATING COMPLEXITY: Dynamic environment: In a competitive ecosystem of continually diversifying life forms, the ecosystem gets more complex, so competing in the ecosystem both enables and requires more complexity.
The exponential increase in complexity produces qualitatively new modes of complexity leveraging beyond initial resources: such as specialization, food chains, parasitical strategies, mutual or cyclical symbioses, discarded products that become new resources, colonization of new environments and energy sources, flexible behaviors based on conditions, greater utilization of existing environments and resources, cooperation within multi-cell colonies, specialization and reproductive coordination within cell colonies (creatures), communication and coordination between similar and different life forms, tool use, tool creation, environment shaping, anticipation and planning, curiosity driven learning, aggregation and recombination of knowledge, resource trading systems, systems to promote positive sum interactions, and suppress negative sum interactions, engineering, invention, science, automation, etc.
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TLDR: Non-trivial environments provide initial opportunities for complexity to improve efficiency. Complexity feedback in ecosystems exponentially accelerates further complexity. Exponential growth of life's complexity on Earth shows no signs of relenting.
Qualitatively new forms of complexity keep appearing. Conscious intelligence, culture, technology and automation are more continuations than breaks from this trend.