I was at Google in ‘06-07, and again ‘09-11, and already there were obvious differences. You just can’t scale “total internal transparency” when the company doubles in size every year.
And at some point you need just plain-old-“good” engineers to make the A/B tests happen and all the other stuff that doesn’t excite the PhDs. And at some point you need product people too, because you start to build products you’re not in the target audience for. And at some point your employees aren’t all going to be fabulously wealthy from an upcoming IPO, and so they’ll start playing political games for coveted titles and $1M+ comp so they too can have their house in Tahoe.
At some point the real world just gets in the way.