Purkinje neurons are found in mammals as well. Huge branches and a lots of input. This paragraph is odd.
This elaborate structures of neurons is a big reason why many neuroscientist don't like point neurons artitecture. Personally I don't think this structure can be approximated by a later or point neurons -- at least not without a significant cost. After reading "More Is Different" I am more open to the idea that substrate matter quite a lot.