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The cease and desist - albeit invalid - and other legal communication was sent privately to the OP. I don't think that counts as defamation.

IANAL, but since this was a C&D notice sent in private and Facebook never made a public announcement regarding it, I don't think it's defamation.

You may have a cda section 230 shield since you were merely a user of the comment section, but you are being treated as the speaker of the comment that is their evidence. It's tenuous but worth a try.

The ACLU loves CDA cases since under CA law, it's loser pay (SLAPP).

Yeah, if I were that guy I'd be playing this up for all it's worth. They're demanding that he prove it's not him, and saying that he "sound(s) guilty"? Is that a joke? If they really said that, that law firm just stepped in it big time.