Worker bees. Not everyone needs to reproduce for the group to prosper.
For the latter part, I'd argue along the lines of the selfish genes (Dawkins): Not an organism wants to reproduce but rather the individual genes. They use the body as a vehicle for that. So it can still make sense for an individual to not reproduce and increase the chances of the same genes to be passed on (e.g., when an uncle without kids supports the kids of his siblings).