More like "make demands without considering the other person at all vs. think for a second about not imposing yourself on other people unnecessarily."
But mostly this article is about the virtues of being a clear communicator and having decent interpersonal skills, which is neither an "ask" nor a "guess" thing.
From my experience, I would assume the one not considering the other person is the one in guess culture, assuming the other person can and ought to read their mind, and the one trying not to impose is the one who actually asks the other person for their opinion or consent, but I can equally see it the other way around, and think the opposite might actually be how you meant it.
I ask all the time! And I'm totally comfortable with "no." But I try to consider the other person first because I think making unreasonable requests repeatedly, which is the subtext of their description of an "asker," blows social capital and just bugs people.
It's not like people in ask cultures just go around asking random strangers to hand over all their money...
Guess also assumes a very similar shared context and understanding. This leads to xenophobia, because "those foreigners" seem so rude simply because they "guess" with a different contexts. Again, asking is superior.