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Based on that, Lemmy seems to be the most reasonable alternative. Can't believe collapsible comments is so rare, that's the main thing (aside from voting) making gigantic comment chains readable.

Isn't Raddle open source? The footer links to https://postmill.xyz/ https://raddle.me/wiki/why_raddle

Oh interesting, I completely missed this. I saw the footer link I thought it was just to the company that runs it.

The more useful thing to users would be a list of alternatives for each closed subreddit. No reddit alternative is going to grow to anything useful from this.

Why not? Typically users of reddit are subscribed to more than one subreddit, so migration to a single destination that can handle all of those communities seems like an obvious choice. The network effect of many users advocating that their second favourite community migrate over to the "new reddit" seems reasonable once people start to establish a userbase there.